Wednesday 10 November 1999, Kelvin Conference Centre, University of Glasgow.

Glasgow 99 Programme

MICROSCOPY OF THE UNDERWORLD - Adam Curtis, Institute of Biomedical and Life Sciences, University of Glasgow.
 
THE USE OF MICROSCOPY-BASED TECHNIQUES TO INVESTIGATE PLANT CELL WALL ARCHITECTURE - Mazz Marry1, I. Max Huxam2, Laurence Tetley2 and Michael C. Jarvis1, 1 Agricultural, Food and Environmental Chemistry, University of Glasgow. 2 Integrated Microscopy Facility, Division of Infection and Immunity, Institute of Biomedical and Life Sciences, University of Glasgow.
 
USING ELECTRONS TO EXPLORE MATERIALS - Alan Craven, Department of Physics and Astronomy, University of Glasgow.
 
EELS INVESTIGATIONS OF THE STRUCTURE AND ELECTRONIC PROPERTIES OF CHROMIUM SPINELS - F. T. Docherty*§, A. J. Craven§ and D. W. McComb*, *Dept. of Chemistry, University of Glasgow. §Dept. of Physics and Astronomy, University of Glasgow.
 
EGGSHELL TAPPING - Sally E. Solomon, Poultry Research Unit, Department of Preclinical Veterinary Studies, University of Glasgow Veterinary School.
 
STUDY OF THE COUPLING BETWEEN SEROTONINERGIC AND NORADRENERGIC NEURONES AND GAMMA MOTONEURONES IN THE CAT - A. Sahal, M.H.Gladden, D. J. Maxwell and E Jankowska, Institute of Biomedical and Life Sciences, West Medical Building, University of Glasgow.
 
ACQUISITION OF DIGITAL DATA IN THE MODERN TEM - Timon F. Fliervoet, Max T. Otten, Wim M. Busing and Marc H. Storms, FEI I Philips Electron Optics, Eindhoven, The Netherlands.
 
COMPUTER ASSISTED QUANTITATIVE ANALYSIS OF AXONAL CYTOSKELETAL RESPONSES TO NON-DISRUPTIVE AXONAL INJURY - W. L. Maxwell and M. Nielson, Laboratory of Human Anatomy, University of Glasgow.

 MICROSCOPY OF THE UNDERWORLD

Adam Curtis,Institute of Biomedical and Life Sciences, University of Glasgow.

The contact between cell and cell and between cell and a material substrate is the region in which adhesions form and are maintained. Though very useful information can be obtained from TEM direct observation of this zone in life is difficult. I shall describe three optical methods, which can be used to examine this region when the cells are grown on surfaces whose refractive index differs appreciably from that of the medium or when appropriate fluorescent labelling can be used. These methods are Interference Reflection Microscopy (IRM), Surface Plasmon Resonance Microscopy (SPRM) and Fluorescence Energy Transfer Microscopy (FETM). Each of these methods has a resolution in the z-axis of at least 10nm and FETM in fact yields no signal when the cell is more than 6nm from the surface. IRM is also a very sensitive method for detecting surface contamination. I shall describe the methods, how they work and how they may be set up and something of the information they give on this nearly two-dimensional world under the cell.

THE USE OF MICROSCOPY-BASED TECHNIQUES TO INVESTIGATE PLANT CELL WALL ARCHITECTURE

Mazz Marry1, I. Max Huxam2, Laurence Tetley2 and Michael C. Jarvis1,
1 Agricultural, Food and Environmental Chemistry, University of Glasgow, Glasgow, G12 8QQ,
2 Integrated Microscopy Facility, Division of Infection and Immunity, Institute of Biomedical and Life Sciences, University of Glasgow, Glasgow, G12 8QQ.

The cell wall of higher plants is composed of at least two major structurally independent but interacting networks. The cellulose-hemicellulose network (about 50% of the wall mass), is embedded in a second network of matrix pectic polysaccharides (about 30% of the total mass). There are many types of cross-links involved in holding carbohydrate moieties within the plant cell wall, all of which directly influence cell-cell adhesion and the mechanical strength of the cell wall. Various techniques can be used to analyse the nature and distribution of the carbohydrate polymers present in plant cell walls. Three such techniques, Fourier Transform Infra-red (FTIR) microspectroscopy, immunogold labelling by TEM and Electron Energy Loss Spectroscopy (EELS) were used to investigate the cell walls from mature sugar beet (Beta vulgaris L. Aztec). Unextracted cell walls and the wall residue remaining following four sequential extractions with a calcium chelator (imidazole) and two additional extractions with sodium carbonate (to remove ester bonds) were studied.

Unlike most dicots, sugar beet have significant levels of ferulic acid in their primary walls. These form ester cross-links, which may strengthen the cell wall. FTIR analysis indicates that the wall residue following imidazole extraction contains a high proportion of both saturated and unsaturated esters, which are removed from the wall by the additional treatment of sodium carbonate. Immuno-gold labelling has provided information on the distribution of both unesterified and methyl-esterified pectic epitopes within the cell wall residue. Successive imidazole extracts removed unesterified pectic epitopes throughout the cell wall, but not from the middle lamella. Elemental maps from the unextracted wall material clearly show calcium ions throughout the wall, with a greater concentration in the middle lamella. It has been known for some time that, in mature tissues, the middle lamella glues together the walls of contiguous cells and forms a separate domain within the wall which mainly consists of calcium-cross-linked pectic polysaccharides, polymers which are rich in galacturonic acid. The EELS-imaging data of the wall residue following imidazole extractions indicate that the unesterified pectic epitopes still present within the middle lamella are not cross-linked by calcium ions.

USING ELECTRONS TO EXPLORE MATERIALS

Alan Craven, Department of Physics and Astronomy, University of Glasgow, Glasgow.G12 8QQ.

Electrons have a number of advantages when it comes to using them to study materials. On the optical side, it is relatively easy to make high brightness sources of highly monochromatic electrons in the laboratory. These electrons can be focused with simple magnetic lenses, allowing the formation of either a high-resolution image of the electrons leaving a thin specimen, as in the transmission electron microscope, or a very small electron probe, as in the case of the scanning electron microscope. On the interaction side, the electrons are scattered quite strongly both elastically and inelastically, leading to a wide range of signals that can provide information about the materials being studied. The skill of the electron microscopist comes in making use of these signals to provide the information sought about the material. The talk will show how such signals can be used to study things like paints, hard wear-resistant coatings, pipeline steels and magnetic devices.


EELS INVESTIGATIONS OF THE STRUCTURE AND ELECTRONIC PROPERTIES OF CHROMIUM SPINELS

F. T. Docherty*§, A. J. Craven§ and D. W. McComb*,
*Dept. of Chemistry, University of Glasgow, Glasgow, G12 8QQ
§Dept. of Physics and Astronomy, University of Glasgow, Glasgow G12 8QQ.
E-mail: francesd@chem.gla.ac.uk

Electron energy loss spectroscopy (EELS) provides information about the energy loss suffered by primary electrons when they interact with a specimen. The resulting energy loss spectrum contains edges due to the excitation of core electrons to unoccupied states in the conduction band. These edges exhibit fine structure, which is dependent on the local chemical and structural environment of the absorbing atom. The understanding of this technique can be improved by examining data collected from a series of compounds with the same crystal structures, to investigate the effect on the fine structure of changing the element in a specific crystallographic site.

Spinels are a suitable class of compounds for a study of this type. The study of a series of chromium spinels, [A]tet[Cr2]octO4, (A=Mg2+, Co2+, Ni2+, Cu2+, Zn2+) will determine the effect on the Cr and O edges of occupancy of the d-orbitals on the neighbouring sites. By correlating the experimental data with electronic structure calculations, we aim to develop an interpretative framework, which can be used to determine the chemistry of more complex materials, improving the fundamental understanding of ELNES.

EGGSHELL TAPPING

Sally E. Solomon, Poultry Research Unit, Department of Preclinical Veterinary Studies, University of Glasgow Veterinary School, Bearsden, Glasgow G61 1QH.

Technology leads knowledge? and in the case of the avian ovum, suspended in its protective albumen sac and encased in a bioceramic shell, the egg, as we know it, has been the testing ground for both traditional and space age methodology.

This paper deals primarily with shell construction and to that end egg white is cited as the foundation layer upon and around which subsequent layers will deposit. Its degree of stiffness is therefore of paramount importance. Low Resolution Nuclear Magnetic Resonance is used to monitor the observed changes in albumen viscosity by measuring its water proton longitudinal relaxation rate.

Light, scanning and transmission microscopy have all had a role to play in resolving the morphology of the eggshell proper and scanning microscopy in particular has enabled the characterisation of the twelve major structural variations which occur in the basal layer of the calcitic eggshell and which in turn influence the direction of further crystal growth The Proscan 1000 non-contact surface profiler provides a rapid and objective analysis of the contours of the inner surface of the eggshell.

A non-invasive measurement of egg quality is now possible using acoustic resonance (Egg Tapping). The paper will illustrate its use and limitations.


STUDY OF THE COUPLING BETWEEN SEROTONINERGIC AND NORADRENERGIC NEURONES AND GAMMA MOTONEURONES IN THE CAT

A. Sahal, M.H.Gladden, D. J. Maxwell and E Jankowska,
Institute of Biomedical and Life Sciences, West Medical Building, University of Glasgow, Glasgow G12 8QQ.

The modualtory effects of serotonin are facilitatory on both a- and g?motoneurones. In contrast, noradrenaline has facilitatory effects on a?motoneurones but depressive effects on g-motoneurones. Since these differences might be reflected in the coupling between descending noradrenergic and serotoninergic nerve fibres with g-motoneurones the coupling was investigated.

Eight g-motoneurones were labelled with rhodamine-dextran injected intracellularly in anaesthetised cats. Serotoninergic and noradrenergic axons were identified with antibodies against dopamine b-hydroxylase and serotonin. Appositions between labelled axons and somata and dendrites were located with three colour confocal laser scanning microscopy by examining series of optical sections gathered at 1mm intervals in 50mm thick sagital sections of the spinal cord.

Serotoninergic and noradrenergic varicosities were found in apposition and close to g-motoneurones. Serotoninergic varicosities formed 1.05 ± 0.10 per 100mm2 with the somata and 14.5 ± 1.95, 11.3 ± 1.09 and 7.4 ± 2.29 appositions per mm length of dendrite within 100, 100-300mm and >300mm distances from the soma, respectively.

The number of appositions of noradrenergic varicosities were 0.6 ± 0.10 per 100mm 2 with somata and 9.2 ± 1.83, 7.6 ± 1.09 and 3.6 ± 2.37 per 100mm length of the dendrites 100, 100-300 and >300mm from the soma respectively.

No major differences were found in the coupling between serotoninergic fibres compared with noradrenergic fibres and g-motoneurones except that the density of serotoninergic appositions was higher. Comparison with data available for coupling between serotoninergic fibres and a-motoneurones suggests that it is similar for a- and g-motoneurones. Comparison of the numbers of varicosities in apposition to those in the immediate surround (in a 5mm shell) suggests that the varicosities are preferentially located on the surface of the g-motoneurones.

 

ACQUISITION OF DIGITAL DATA IN THE MODERN TEM

Timon F. Fliervoet, Max T. Otten, Wim M. Busing and Marc H. Storms, FEI I Philips Electron Optics, Eindhoven, The Netherlands.
tff@ffi-ni.feico.com

A thorough characterisation of a material is the start of a fundamental understanding of the material's properties. Morphology, crystal structure, chemical composition, interface structure, surfaces and defects all have their influence on the properties of materials. In (advanced) materials science and design there is an increasing need for the determination of the chemical and electronic structure of materials. As microstructures become more and more complex, the dimensions to be analysed are becoming smaller. Transmission Electron Microscopy (TEM) has proven to be a very powerful technique for studying a range of materials down to sub-nanometer levels. It generates a wide range of signals, carrying different types of valuable information. TEM also contributed significantly in finding answers to fundamental questions about the functionality of ongoing physiological and pathological processes.

However, in the past TEMs, equipped with many detectors, were surrounded by a single proliferation with PC's, monitors and mice. These add-on monitors and computers often caused stray-fields and interferences, and require too much room. Furthermore, it was difficult to operate the different computers with different operating systems. Most important, the interaction between microscopy and the data was lost.

The Tecnai series of TEM has been especially designed to acquire and process these digital signals efficiently and simultaneously. All the detectors - including STEM, TV and CCD cameras, EDX and EELS detectors - are fully embedded into one Tecnai operating system, with one monitor, mouse and keyboard. Tecnai, the modem TEM, makes microscopy easier then ever.


COMPUTER ASSISTED QUANTITATIVE ANALYSIS OF AXONAL CYTOSKELETAL RESPONSES TO NON-DISRUPTIVE AXONAL INJURY

W. L. Maxwell and M. Nielson, Laboratory of Human Anatomy, University of Glasgow, Glasgow G12 8QQ.

Axons within the white matter of the brain are exposed to transient tensile strain during diffuse axonal injury (DAI) which is the commonest cause of death in children, and adults under the age of 40. A great deal of attention in a number of experimental laboratories over the last 15 years has been focused upon the characterisation of axonal pathology with the aim of defining the pathological mechanisms operating in such models of DAI. It is clear that axons are not fragmented at the time of injury but rather enter a so-called "pathological cascade" of events which lead to axotomy over hours and days after injury. A Kontron Elektronik-Vidas Image Analysis has been used to develop a program for the quantitative analysis of responses by components of the axonal cytoskeleton, microtubules and neurofilaments, to the application of transient mechanical strain.

The methodology used in this study will be described. We shall also demonstrate that microtubules and neurofilaments respond with different rates to injury, that cytoskeletal responses can be quantified, that different sizes of axons respond in different ways to such injury and that there is a potential for the use of post-traumatic therapy to ameliorate the effects of injury.