Professor Paul Thompson
Chair in Zoology
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01381 600841
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lighthouse@abdn.ac.uk
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Lighthouse Field Station, George Street, Cromarty, Ross-shire, IV11 8YJ
Chair in Zoology
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Personal Details
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Biography
1982, B.Sc. Biology, University of York
1987, PhD Zoology, University of Aberdeen
1990, Established the University’s Lighthouse Field Station
2005, Appointed to a Personal Chair in Zoology
2011, Elected Fellow of the Royal Society of Edinburgh (RSE)
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Research Interests
Current research aims to assess how natural and anthropogenic environmental variations influence the behaviour, physiology and dynamics of marine mammal and seabird populations. These questions have been approached by conducting long-term and comparative studies of key populations; primarily in Scottish waters but including some work overseas. These studies have been used to address a range of applied and theoretical questions, often requiring an inter-disciplinary approach and regularly involving collaboration with other research groups both in the UK and abroad. Topics of particular interest have included interactions between wildlife populations and fisheries, the impact of anthropogenic noise on marine mammal biology, seal foraging and breeding strategies, and effects of changing prey stocks and climate change on the population dynamics of marine top predators.
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Research Grants
2011-2012, Marine Scotland, Assessment of methods for monitoring marine mammals at marine renewable energy developments
2010-2012, Moray Offshore Renewables Ltd & Beatrice Offshore Wind Ltd, Development of a framework for assessing the impact of windfarm construction marine mammal populations.
2009-2013, DECC, Assessing the impact of seismic survey noise on cetaceans
2005-2008, SEERAD, Distribution and abundance of bottlenose dolphins in Scottish waters.
2005-2007, EU Framework 6, DOWNVIND - Assessing the impact of offshore wind turbines on seabirds and cetaceans.
2004-2011, Scottish Natural Heritage, Site condition monitoring for the Moray Firth Special Area of Conservation
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External Responsibilities
Member of Marine Scotland Science Advisory Board
University of Aberdeen representative on the University of the Highlands & Islands Research Committee
Member of the US National Research Council Committee on Ecological Effects of Mariculture
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Lighthouse Field Station
Paul Thompson is currently Director of the Lighthouse Field Station, which provides a base for the University's research and teaching in the Moray Firth. Further details of the Field Station's staff, research and training programmes can be found be found at www.abdn.ac.uk/zoology/lighthouse.
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Publications
Contributions to Journals
Articles
- Cheney, BJ., Thompson, PM., Ingram, SN., Hammond, PS., Stevick, PT., Durban, JW., Culloch, RM., Elwen, SH., Mandleberg, L., Janik, VM., Quick, NJ., Islas-Villanueva, V., Robinson, KP., Costa, M., Eisfeld, SM., Walters, A., Phillips, C., Weir, CR., Evans, PGH., Anderwald, P., Reid, RJ., Reid, JB. & Wilson, B. (2013). 'Integrating multiple data sources to assess the distribution and abundance of bottlenose dolphins Tursiops truncatus in Scottish waters'. Mammal Review, vol 43, no. 1, pp. 71-88.
[Online] DOI: 10.1111/j.1365-2907.2011.00208.x - Bailey, HR., Corkrey, R., Cheney, BJ. & Thompson, PM. (2013). 'Analyzing temporally correlated dolphin sightings data using generalized estimating equations'. Marine Mammal Science, vol 29, no. 1, pp. 123-141.
[Online] DOI: 10.1111/j.1748-7692.2011.00552.x - Cordes, L., Duck, C., MacKey, BL., Hall, A. & Thompson, PM. (2011). 'Long-term patterns in harbour seal site-use and the consequences for managing protected areas'. Animal Conservation, vol 14, no. 4, pp. 430-438.
[Online] DOI: 10.1111/j.1469-1795.2011.00445.x - Orzack, SH., Steiner, UK., Tuljapurkar, S. & Thompson, PM. (2011). 'Static and dynamic expression of life history traits in the Northern Fulmar (Fulmarus glacialis)'. Oikos, vol 120, no. 3, pp. 369-380.
[Online] DOI: 10.1111/j.1600-0706.2010.17996.x - Lonergan, M., Hall, A., Thompson, H., Thompson, PM., Pomeroy, P. & Harwood, J. (2010). 'Comparison of the 1988 and 2002 phocine distemper epizootics in British harbour seal Phoca vitulina populations'. Diseases of Aquatic Organisms, vol 88, no. 3, pp. 183-188.
[Online] DOI: 10.3354/dao02153 - Bailey, HR., Clay, G., Coates, EA., Lusseau, D., Senior, B. & Thompson, PM. (2010). 'Using T-PODs to assess variations in the occurrence of coastal bottlenose dolphins and harbour porpoises'. Aquatic Conservation: Marine and Freshwater Ecosystems, vol 20, no. 2, pp. 150-158.
[Online] DOI: 10.1002/aqc.1060 - Bailey, HR. & Thompson, PM. (2010). 'The effect of oceanographic features on the fine-scale foraging movements of bottlenose dolphins'. Marine Ecology Progress Series, vol 418, pp. 223-233.
[Online] DOI: 10.3354/meps08789 - Thompson, PM., Lusseau, D., Barton, TR., Simmons, D., Rusin, J. & Bailey, HR. (2010). 'Assessing the responses of coastal cetaceans to the construction of offshore wind turbines'. Marine Pollution Bulletin, vol 60, no. 8, pp. 1200-1208.
[Online] DOI: 10.1016/j.marpolbul.2010.03.030 - Bailey, HR., Senior, BL., Simmons, D., Rusin, J., Picken, G. & Thompson, PM. (2010). 'Assessing underwater noise levels during pile-driving at an offshore windfarm and its potential effects on marine mammals'. Marine Pollution Bulletin, vol 60, no. 6, pp. 888-897.
[Online] DOI: 10.1016/j.marpolbul.2010.01.003 - Bailey, HR. & Thompson, PM. (2009). 'Using marine mammal habitat modelling to identify priority conservation zones within a marine protected area'. Marine Ecology Progress Series, vol 378, pp. 279-287.
[Online] DOI: 10.3354/meps07887 - Lewis, S., Elston, D., Daunt, F., Cheney, BJ. & Thompson, PM. (2009). 'Effects of extrinsic and intrinsic factors on breeding success in a long lived seabird'. Oikos, vol 118, no. 4, pp. 521-528.
[Online] DOI: 10.1111/j.1600-0706.2008.17308.x - Corkrey, R., Brooks, S., Lusseau, D., Parsons, K., Durban, JW., Hammond, PS. & Thompson, PM. (2008). 'A Bayesian Capture-Recapture Population Model With Simultaneous Estimation of Heterogeneity'. Journal of the American Statistical Association, vol 103, no. 483, pp. 948-960.
[Online] DOI: 10.1198/016214507000001256 - Butler, JRA., Middlemas, SJ., McKelvey, SA., McMyn, I., Leyshon, B., Walker, I., Thompson, PM., Boyd, IL., Duck, C., Armstrong, JD., Graham, IM. & Baxter, JM. (2008). 'The Moray Firth Seal Management Plan: an adaptive framework for balancing the conservation of seals, salmon, fisheries and wildlife tourism in the UK'. Aquatic Conservation: Marine and Freshwater Ecosystems, vol 18, pp. 1025-1038.
[Online] DOI: 10.1002/aqc.923 - Mallory, ML., Gaston, AJ., Forbes, MR., Gilchrist, HG., Cheney, BJ., Lewis, S. & Thompson, PM. (2008). 'Flexible incubation rhythm in northern fulmars: a comparison between oceanographic zones'. Marine Biology, vol 154, no. 6, pp. 1031-1040.
[Online] DOI: 10.1007/s00227-008-0994-z - Thompson, PM. & Wheeler, H. (2008). 'Photo-ID-based estimates of reproductive patterns in female harbor seals'. Marine Mammal Science, vol 24, no. 1, pp. 138-146.
[Online] DOI: 10.1111/j.1748-7692.2007.00179.x - Jones, OR., Gaillard, J-M, Tuljapurkar, S., Alho, JS., Armitage, KB., Becker, PH., Bize, P., Brommer, J., Charmantier, A., Charpentier, M., Clutton-Brock, T., Dobson, FS., Festa-Bianchet, M., Gustafsson, L., Jensen, H., Jones, CG., Lillandt, B-G, McCleery, R., Merila, J., Neuhaus, P., Nicoll, MAC., Norris, K., Oli, MK., Pemberton, J., Pietiainen, H., Ringsby, TH., Roulin, A., Saether, B-E, Setchell, JM., Sheldon, BC., Thompson, PM., Weimerskirch, H., Wickings, EJ. & Coulson, T. (2008). 'Senescence rates are determined by ranking on the fast-slow life-history continuum'. Ecology Letters, vol 11, no. 7, pp. 664-673.
[Online] DOI: 10.1111/j.1461-0248.2008.01187.x - MacKey, BL., Durban, J., Middlemas, S. & Thompson, PM. (2008). 'A Bayesian estimate of harbour seal survival using sparse photo-identification data'. Journal of Zoology, vol 274, no. 1, pp. 18-27.
[Online] DOI: 10.1111/j.1469-7998.2007.00352.x - Thompson, PM., MacKey, BL., Barton, TR., Duck, C. & Butler, JRA. (2007). 'Assessing the potential impact of salmon fisheries management on the conservation status of harbour seals (Phoca vitulina) in north-east Scotland'. Animal Conservation, vol 10, no. 1, pp. 48-56.
[Online] DOI: 10.1111/j.1469-1795.2006.00066.x - Baker, JD. & Thompson, PM. (2007). 'Temporal and spatial variation in age-specific survival rates of a long-lived mammal, the Hawaiian monk seal'. Proceedings of the Royal Society of London. B, Biological Sciences, vol 274, no. 1608, pp. 407-415.
[Online] DOI: 10.1098/rspb.2006.3737 - Hastie, GD., Wilson, B. & Thompson, PM. (2006). 'Diving deep in a foraging hotspot: acoustic insights into bottlenose dolphin dive depths and feeding behaviour'. Marine Biology, vol 148, no. 5, pp. 1181-1188.
[Online] DOI: 10.1007/s00227-005-0143-x - Lusseau, D., Wilson, B., Hammond, PS., Grellier, K., Durban, J., Parsons, KM., Barton, TR. & Thompson, PM. (2006). 'Quantifying the influence of sociality on population structure in bottlenose dolphins'. Journal of Animal Ecology, vol 75, no. 1, pp. 14-24.
[Online] DOI: 10.1111/j.1365-2656.2005.01013.x - Bailey, HR. & Thompson, PM. (2006). 'Quantitative analysis of bottlenose dolphin movement patterns and their relationship with foraging'. Journal of Animal Ecology, vol 75, no. 2, pp. 456-465.
[Online] DOI: 10.1111/j.1365-2656.2006.01066.x - Butler, JRA., Middlemas, SJ., Graham, IM., Thompson, PM. & Armstrong, JD. (2006). 'Modelling the impacts of removing seal predation from Atlantic salmon, Salmo salar, rivers in Scotland: a tool for targeting conflict resolution'. Fisheries Management and Ecology, vol 13, no. 5, pp. 285-291.
[Online] DOI: 10.1111/j.1365-2400.2006.00504.x - Harkonen, T., Dietz, R., Reijnders, P., Teilmann, J., Harding, K., Hall, A., Brasseur, S., Siebert, U., Goodman, SJ., Jepson, PD., Rasmussen, TD. & Thompson, PM. (2006). 'The 1988 and 2002 phocine distemper virus epidemics in European harbour seals'. Diseases of Aquatic Organisms, vol 68, no. 2, pp. 115-130.
- Thompson, PM. (2006). 'Identifying drivers of change'. Conservation Biology, no. 12, pp. 143-156.
- Hastie, GD., Swift, RJ., Slesser, G., Thompson, PM. & Turrell, W. (2005). 'Environmental models for predicting oceanic dolphin habitat in the northeast Atlantic'. ICES Journal of Marine Science, vol 62, pp. 760-770.
[Online] DOI: 10.1016/j.icesjms.2005.02.004 - Durban, J., Elston, D., Ellifrit, DK., Dickson, E., Hammond, PS. & Thompson, PM. (2005). 'Multi-site mark-recapture population estimates with Bayesian model determination'. Marine Mammal Science, vol 21, pp. 80-92.
[Online] DOI: 10.1111/j.1748-7692.2005.tb01209.x - Grosbois, V. & Thompson, PM. (2005). 'North Atlantic climate variation influences survival in adult fulmars'. Oikos, vol 109, no. 2, pp. 273-290.
[Online] DOI: 10.1111/j.0030-1299.2005.13774.x - Middlemas, S., Barton, TR., Armstrong, JD. & Thompson, PM. (2005). 'Functional and aggregative responses of harbour seals to changes in salmonid abundance'. Proceedings of the Royal Society of London. B, Biological Sciences, vol 273, no. 1583, pp. 193-198.
[Online] DOI: 10.1098/rspb.2005.3215 - Foster, G., Holmes, B., Steigerwalt, AG., Lawson, PA., Thorne, P., Byrer, DE., Ross, HM., Xerry, J., Thompson, PM. & Collins, MD. (2004). 'Camphylobacter insulaenigrae sp. nov. Isolated from marine mammals'. International Journal of Systematic and Evolutionary Microbiology, vol 54, pp. 2369-2373.
[Online] DOI: 10.1099/ijs.0.63147-0 - Lusseau, D., Wilson, B., Grellier, K., Barton, TR., Thompson, PM., Williams, R. & Hammond, PS. (2004). 'Parallel influence of climate on the behaviour of Pacific killer whales and Atlantic bottlenose dolphins'. Ecology Letters, vol 7, no. 11, pp. 1068-1076.
[Online] DOI: 10.1111/j.1461-0248.2004.00669.x - Hastie, GD., Wilson, B., Wilson, L., Parsons, KM. & Thompson, PM. (2004). 'Functional mechanisms underlying cetacean distribution patterns: hotspots for bottlenose dolphins are linked to foraging'. Marine Biology, vol 144, pp. 397-403.
[Online] DOI: 10.1007/s00227-003-1195-4 - Wilson, B., Reid, RJ., Grellier, K., Thompson, PM. & Hammond, PS. (2004). 'Considering the temporal when managing the spatial: a population range expansion impacts protected areas based management from bottlenose dolphins'. Animal Conservation, vol 7, no. 4, pp. 331-338.
[Online] DOI: 10.1017/S1367943004001581 - Thompson, PM., White, S. & Dickson, E. (2004). 'Co-variation in the probabilities of sighting harbour porpoises and bottlenose dolphins'. Marine Mammal Science, vol 20, pp. 322-328.
[Online] DOI: 10.1111/j.1748-7692.2004.tb01160.x - Michel, P., Ollason, JG., Grosbois, V. & Thompson, PM. (2003). 'The influence of body size, breeding experience and evironmental variability on egg size in the northern fulmar ( Fulmarus glacialis )'. Journal of Zoology, vol 261, pp. 427-432.
[Online] DOI: 10.1017/S0952836903004291 - Hastie, GD., Barton, TR., Swift, RJ., Grellier, K., Hammond, PS. & Thompson, PM. (2003). 'Distribution of small cetaceans within a candidate Special Area of Conservation; implications for management'. Journal of Cetacean Research and Management, vol 5, pp. 261-266.
- Hastie, GD. & Thompson, PM. (2003). 'Bottlenose Dolphins increase breathing synchrony in response to boat traffic'. Marine Mammal Science, vol 19, no. 1, pp. 74-84.
[Online] DOI: 10.1111/j.1748-7692.2003.tb01093.x - Van Parijs, SM., Corkeron, PJ., Harvey, J., Hayes, S., Mellinger, D., Rouget, P., Thompson, PM., Wahlberg, M. & Kovacs, KM. (2003). 'Patterns in the vocalizations of male harbor seals'. Journal of the Acoustical Society of America, vol 113, no. 6, pp. 3403-3410.
[Online] DOI: 10.1121/1.1568943 - Hastie, GD., Wilson, B. & Thompson, PM. (2003). 'Fine-scale habitat selection by coastal bottlenose dolphins: application of a new land-based video-montage technique'. Canadian Journal Of Zoology/Revue Canadien De Zoologie, vol 81, pp. 469-478.
[Online] DOI: 10.1139/z03-028 - Michel, P. & Thompson, PM. (2003). 'A simple photogrammetric technique for estimating egg volume from field measurements'. Atlantic Seabirds, vol 5, no. 1, pp. 31-34.
- Parsons, KM., Durban, J., Claridge, DE., Balcombe, KC., Noble, LR. & Thompson, PM. (2003). 'Kinship as a basis for alliance formation between male bottlenose dolphins, Tursiops truncatus , in the Bahamas'. Animal Behaviour, vol 66, pp. 185-194.
[Online] DOI: 10.1006/anbe.2003.2186 - Grellier, K., Hammond, PS., Wilson, B., Sanders-Reed, C. & Thompson, PM. (2003). 'Use of photo-identification data to quantify mother-calf association patterns in bottlenose dolphins'. Canadian Journal Of Zoology/Revue Canadien De Zoologie, vol 81, pp. 1421-1427.
[Online] DOI: 10.1139/z03-132 - Thompson, PM., Thompson, H. & Hall, AJ. (2002). 'Prevalence of morbillivirus antibodies in Scottish harbour seals'. Veterinary Record, vol 151, pp. 609-610.
- Mendes, S., Turrell, W., Lutkebohle, T. & Thompson, PM. (2002). 'Influence of the tidal cycle and a tidal intrusion front on the spatio-temporal distribution of coastal bottlenose dolphins'. Marine Ecology Progress Series, vol 239, pp. 221-229.
- Parsons, KM., Noble, LR., Reid, RJ. & Thompson, PM. (2002). 'Mitochondrial genetic diversity and population structuring of UK bottlenose dolphins ( Tursiops truncatus ): is the NE Scotland Population demographically and geographically isolated?'. Biological Conservation, vol 108, pp. 175-182.
[Online] DOI: 10.1016/S0006-3207(02)00103-9 - Denardo, C., Hastie, G., Leaper, R., Wilson, B. & Thompson, PM. (2001). 'A new technique to measure spatial relationships within groups of free-ranging coastal cetaceans'. Journal of Applied Ecology, vol 38, no. 4, pp. 888-895.
[Online] DOI: 10.1046/j.1365-2664.2001.00639.x - Thompson, PM., Van Parijs, S. & Kovacs, KM. (2001). 'Local declines in the abundance of harbour seals:implications for the designation and monitoring of protected areas'. Journal of Applied Ecology, vol 38, pp. 117-125.
[Online] DOI: 10.1046/j.1365-2664.2001.00571.x - Thompson, PM. & Ollason, JG. (2001). 'Lagged effects of ocean climate change on fulmar population dynamics'. Nature, vol 413, pp. 417-420.
[Online] DOI: 10.1038/35096558 - Van Parijs, SM., Janik, VM. & Thompson, PM. (2000). 'Display-area size, tenure length, and site fidelity in the aquatically mating male harbour seal, Phoca vitulina'. Canadian Journal Of Zoology/Revue Canadien De Zoologie, vol 78, no. 12, pp. 2209-2217.
[Online] DOI: 10.1139/z00-165 - Wilson, B., Arnold, H., Bearzi, G., Fortuna, CM., Gaspar, R., Ingram, S., Liret, C., Pribanic, S., Read, AJ., Ridoux, V., Schneider, K., Urian, KW., Wells, RS., Wood, C., Thompson, PM. & Hammond, PS. (1999). 'Epidermal diseases in bottlenose dolphins: impacts of natural and anthropogenic factors'. Proceedings of the Royal Society of London. B, Biological Sciences, vol 266, pp. 1077-1083.
- Tollit, DJ., Steward, MJ., Thompson, PM., Pierce, GJ., Santos, MB. & Hughes, S. (1997). 'Species and size differences in the digestion of otoliths and beaks: Implications for estimates of pinniped diet composition'. Canadian Journal of Fisheries and Aquatic Sciences, vol 54, no. 1, pp. 105-119.
- Thompson, PM., Pierce, GJ., Hislop, JRG., Miller, D. & Diack, J. (1991). 'Winter Foraging by Common Seals (Phoca Vitulina) in Relation to Food Availability in the Inner Moray Firth, N.E. Scotland'. Journal of Animal Ecology, vol 60, no. 1, pp. 283-294.
- Pierce, GJ., Thompson, PM., Miller, A., Diack, JSW., Miller, D. & Boyle, PR. (1991). 'Seasonal variation in the diet of common seals (Phoca vitulina) in the Moray Firth area of Scotland'. Journal of Zoology, vol 223, no. 4, pp. 641-652.
[Online] DOI: 10.1111/j.1469-7998.1991.tb04393.x
Letters
- Gales, NJ., Bowen, WD., Johnston, DW., Kovacs, KM., Littnan, CL., Perrin, WF., Reynolds, JE. & Thompson, PM. (2009). 'Guidelines for the treatment of marine mammals in field research'. Marine Mammal Science, vol 25, no. 3, pp. 725-736.
[Online] DOI: 10.1111/j.1748-7692.2008.00279.x
Comments and Debates
- Pierce, GJ., Miller, A., Thompson, PM. & Hislop, JRG. (1991). 'Prey Remains in Gray Seal (Halichoerus-Grypus) Feces from the Moray Firth, North East Scotland'. Journal of Zoology, vol 224, no. 2, pp. 337-341.
Editorials
- Thompson, P., Ingram, SN., Lonergan, M., Northridge, S., Hall, A. & Wilson, B. (2007). 'Climate change causing starvation in harbour porpoises?'. Biology Letters, vol 3, no. 5, pp. 533-535.
[Online] DOI: 10.1098/rsbl.2007.0076 - Thompson, PM. (2007). 'Developing water quality standards for coastal dolphins'. Marine Pollution Bulletin, vol 54, no. 2, pp. 123-127.
[Online] DOI: 10.1016/j.marpolbul.2006.11.026
Chapters in Books, Reports and Conference Proceedings
Chapters
- Durant, JM., Stenseth, NC., Anker-Nilssen, T., Harris, MP., Thompson, PM. & Wanless, S. (2004). 'Marine birds and climate fluctuation in the North Atlantic'. in Marine Ecosystems and Climate Variation: The North Atlantic. Marine Ecosystems and Climate Variation: The North Atlantic, pp. 95-105.
Contributions to Conferences
Posters
- Cheney, BJ., Lusseau, D., Hammond, PS. & Thompson, PM. 'Inflation factors influence the estimation of trends in cetacean abundance'. 19th Biennial Conference on the Biology of Marine Mammals, Tampa, FL, United States, 27/11/11 - 2/12/11,.
- Cheney, BJ., Barton, TR. & Thompson, PM. (2008). 'Using laser metrics to measure wild bottlenose dolphins'. 22nd Annual European Cetacean Society Conference, Egmond aan Zee, Netherlands, 10/03/08 - 12/03/08,.
Books and Reports
Books
- Petersen, C., Costa-Pierce, BA., Dumbauld, BR., Friedman, C., Hoffmann, EE., Kite-Powell, H., Manahan, DT., O'Beirn, F., Paine, RT., Thompson, PM., Whitlach, R. & Committee on Best Practices for Shellfish Mariculture and the Effects of Commercial Activities in Drakes Estero (2010). 'Ecosystem Concepts for Sustainable Bivalve Mariculture'. The National Academies Press, Washington, DC, USA.
- Petersen, CH., Costa-Pierce, BA., Dumbauld, BR., Friedman, C., Hoffmann, EE., Kite-Powell, H., Manahan, DT., O'Beirn, F., Paine, RT., Thompson, PM., Whitlach, R. & Committee on Best Practices for Shellfish Mariculture and the Effects of Commercial Activities in Drakes Estero; National Research Council (2009). 'Shellfish Mariculture in Drakes Estero, Point Reyes National Seashore, California'. The National Academies Press, Washington, DC, USA.
Commissioned Reports
- Cheney, BJ., Corkrey, R., Quick, NJ., Janik, VM., Islas-Villanueva, V., Hammond, PS. & Thompson, PM. (2012). 'Site Condition Monitoring of bottlenose dolphins within the Moray Firth Special Area of Conservation: 2008-2010'. vol. 512, Scottish Natural Heritage.
- Thompson, PM., Cheney, BJ., Ingram, S., Stevick, P., Wilson, B. & Hammond, PS. (2011). 'Distribution, abundance and population structure of bottlenose dolphins in Scottish waters: Scottish Government and Scottish Natural Heritage funded report'. ol. 354, Scottish Natural Heritage.
- Walls, RJ., Pendlebury, CJ., Budgey, R., Brookes, KL. & Thompson, PM. (2009). 'Revised best practice guidance for the use of remote techniques for ornithological monitoring at offshore windfarms'. COWRIE, United Kingdom.
- Senior, BL., Bailey, HR., Lusseau, D., Foote, AD. & Thompson, PM. (2008). 'Anthropogenic noise in the Moray Firth SAC: potential sources and impacts on bottlenose dolphins'. vol. Commissioned Report No 265, Scottish Natural Heritage.
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